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Free radical biology & medicine 42 , 14701480, doi: 10.1016/j.freeradbiomed.2007.01.044 (2007)

GHK-Cu upregulates several tumor suppressor genes (caspases, growth arrest and DNA damage-inducible genes) and downregulates genes associated with metastasis, invasion, and angiogenesis

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showed that the human klotho gene encodes -klotho, a multifunctional protein that includes three types: full-length transmembrane -Klotho, truncated soluble -Klotho, and secreted -Klotho (72)

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