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Serine-hydroxymethyltransferase ( SHMT ), thymidilate synthase ( TYMS ), and dihydrofolate reductase ( DHFR ) SHMT uses serine as the one-carbon donor for the conversion of THF into 5,10-methyleneTHF, that can be used for thymidylate synthesis in the reaction catalyzed by TYMS that produces dTMP and DHF, which is then reduced back to THF by DHFR (Figure 1)

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