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No single direct pharmacological mechanism has been definitively established

active lesions are enriched in collagen-producing cells, aggregates of fibronectin and collagen, mesenchymal perivascular aggregates of platelet-derived growth factor receptor (PDGFR)-bearing cells, and pro-fibrotic factors such as biglycan and decorin

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Tryptophan at position 9 is the principal photolabile residue and oxidises readily, with tyrosine at position 2 and histidine at position 6 secondary sites of concern