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This process, known as glutathione conjugation, is catalysed by enzymes called glutathione S-transferases (GSTs)

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In fact, transient overexpression of EAAT3 in hippocampal HT22 cells led to increased intracellular GSH levels in the presence of high glutamate concentrations, offering protection against oxidative glutamate toxicity [44, 45]

This four electron reduction proceeds in two chemical steps both catalyzed by the same enzyme, dihydrofolate reductase